Under 1stLt Herbert Sobel, Easy Company is shipped to England to prepare for D-Day. As training progresses, Sobel's inadequacies as a leader in the field become more apparent and cause most of the non-commissioned officers in the company to attempt to resign en masse.

Easy Company lands in Normandy but is scattered across the region and away from the designated drop zone. 1stLt Winters takes command. With a small group of men, Winters takes out a set of German gun emplacements at Brecourt, winning the respect of his fellow soldiers.

Easy Company fights in the Battle of Carentan in which they lose several men. Rumours circulate that Lt Speirs killed a group of German POWs. Pvt Albert Blithe is spurred into action by Winters during the Battle of Bloody Gulch but killed days later by a German sniper.

Replacements join Easy Company. The Company parachutes into the Netherlands where they liberate Eindhoven. During combat in Nuenen, Sgt Denver "Bull" Randleman, the replacements' immediate superior, evades German soldiers after being cut off from his unit.

Winters writes a report on the challenge of an unexpected resistance to a German attack, and is haunted by his conscience after shooting a young German SS soldier. Operation Pegasus is depicted. Easy Company is called to Bastogne at the start of the Battle of the Bulge.

Easy Company experiences the Battle of the Bulge, having to hold ground near Bastogne while running low on ammunition and supplies. Medic Eugene "Doc" Roe helps out his fellow soldiers where he can, whilst also scrounging for medical supplies which are dangerously low.

Easy Company battles near Foy, Belgium, losing numerous men. 1stLt Norman Dike is relieved by 1stLt Ronald Speirs. 1stSg Carwood Lipton attempts to keep the morale of the men up as they endure their trials in the forest near Foy, earning him a battlefield promotion.

Easy Company carries out a dangerous mission in Haguenau as David Webster returns from hospital, along with new replacement 2ndLt Jones. Webster eventually reintegrates with the others but their experiences at Bastogne have made them weary and distant towards him.

As Nixon scrounges for his favoured whiskey, Vat 69, Easy Company enters Germany. Some of the men on patrol stumble across a concentration camp near Landsberg and free the prisoners after realising the guards have deserted. The sight of the victims leaves many appalled.

The company captures the Eagle's Nest in Berchtesgaden and discovers Herman Goering's house, then heads out to Austria where the end of the war in Europe is announced. While those with enough points go home, the rest stay behind until the Pacific War is declared over.